Author: (Chang & Castro, 1999)
Comments: Occurs in the main river and major tributaries, always over sandy beaches.  Stomach contents include larvae of aquatic insects, small seeds and debris.  The holotype (male, 11.59 cm SL) carried 53 fertilized eggs (3.2-3.4 mm diameter) on its lips and another specimen was carrying 31 unfertilized eggs (3.1-3.4 mm diameter).