Author: Regan, 1908
FBname: Machaca
Remark: See Howes (1982: 29-30) for remarks on taxonomy, Burcham (1988), Bussing (1993), and Horn (1997) for observations on ecology of the species. Food fish.
Comments: Found in lakes, rivers and streams between 0 and 600 m elevation.  The young feed on terrestrial and aquatic insects, leaves, fruits and seeds but become mainly herbivorous when adult.  Eggs are deposited in nests located in sand bottoms of streams.