Author: (Bonaparte, 1839)
Comments: Found in middle reaches of rivers and main tributaries, in piedmont, with moderate to swift current, over gravel bottom.  Occurs in groups.  Omnivorous, feeding mainly on aquatic insects and epilithic algae.  Undertakes small spawning migration in large groups towards main tributaries. Spawns in swift current on gravel (Ref. 59043).  Reported to spawn in March-May (Ref. 26100) and in May-June (Ref. 59043).  Threatened due to the introduction of other species and habitat destruction (Ref. 26100).