Author: Kim, Jeon & Suk, 2014
Comments: Occurs mostly in clear and shallow (40-120 cm deep) low to medium-gradient large creeks to medium rivers, over coarse sand to pebble substrates.  Inhabits backwaters or shallow pools near vegetation from autumn to spring.  Large individual are usually collected in rock- and gravel-bottomed riffles during summer. The type series was collected together with the congener <i>T. lanceolata, along with the following species in the subfamily Acheilognathinae:  <i>Acheilognathus macropterus, <i>A. majusculus, <i>A. rhombeus, <i>A. yamatsutae, <i>Rhodeus ocellatus, <i>R. notatus and <i>R. sinensis.  Spawns from April to August.  Spawning males aggresively defend a territory around a mussel-spawning site.  Also collected together with unionid mussels (<i>Unio douglasiae, <i>Anodonta woodiana, <i>Lanceolaria grayana and <i>Solenaia triangularis, though the major host mussels are still not known (Ref. 97360).