Author: Vari, Sidlauskas & Le Bail, 2012
Comments: The type locality of the holotype is a side channel of  Litani River that isolated a small, elongate, forested island.  The channel dominantly consists of shallow cataracts over a bedrock substrate strewn with boulders in the upper portions and a small, shallow (less than 2 m deep) backwater over a sand substrate in the downstream portion.  The channel margins  are composed of shallow pockets of slackwater habitat, often with submerged vegetation and shaded by overhanging bank vegetation extending into the water.  Found in shallow (0.50 m deep) tributary to the  main river in an area with relatively rapid water flow (around 1m/s) and principally sandy substrate with some rocky bars and large pieces of dead wood (Ref. 90234).