Author: (Günther, 1880)
Comments: The specimens were recently collected from seven rivers, where the substrate was mainly composed of sandstone boulders with patches of coarse sand and pebbles. It was found in well-oxygenated waters (6.9-11.2 mg/l) with strong current; water turbidity 1.29-75.1 NTU, conductivity 77-4.530 μS/cm, pH 7.8-8.8, and temperature 20.7-31.4°C (Ref. 121236).