Author: (Regan, 1908)
Remark: Reviewed by Bussing (1998: 348), with key, diagnosis, geographical distribution, and figures.
Comments: Inhabits rivers and rivulets of all velocities, slow to fast moving.  Feeds on algae, leaves, fruits, aquatic insects and bottom detritus.  Found up to 1800 feet of elevation.  Spawns in caves and crevices of rocks on river bottoms and produces 200-250 offspring per spawning.