Author: (Steindachner, 1881)
Comments: In Brunei, both adults and juveniles were encountered frequently in small, shallow and clear, tannin-stained mangrove pools and rivulets among <i>Nypa</i> and <i>Rhizophora</i> roots and litter at low tide, usually well back from the main stream (Ref. 56945).  In Indonesia, specimens were also collected from tannin-stained, fresh-tasting water, but a non-mangrove habitat (Ref. 56945).