Author: Greenwood, 1959
Comments: Inhabits areas with firm substrate (sand and rock) (Ref. 4983).  Gut contents showed that this species preys on a variety of invertebrate animals and possibly even small fish.  Predominant food organisms were insects (esp.  dipterous larvae but also Ephemeroptera & Trichoptera larvae).  The non-insect food were fish, oligochaete worms, prawns, plant tissue, ostracods, and fragments of lamellibranchs & gastropod shells (Ref. 558).