Author: Covain & Fisch-Muller, 2012
Comments: This species was encountered immediately upstream of a 30 m high waterfall and was collected downstream of the waterfall.  The area of the river where this species was found was 10 m wide with shallow clear water and a bottom substrate consisting of gravel, pebbles, boulders, bed rock and sand mainly constituted of iron hydroxide (Ref. 90209).