Author: Armbruster & de Souza, 2005
Comments: Occurs among submerged, dead wood in modest to swift flowing water (Ref. 54531). Inhabiting largest part of freshwaters, from headwaters to upstream of the lower falls, with rocky substrate consisting of boulders, stones, gravels, and sand; the exposed wet rocks are also often covered by the Podostemaceae <i>Mourera fluviatilis</i> and <i>Apinagia richardiana</i> (Ref. 90186).