Author: Kotlík, Tsigenopoulos, Ráb & Berrebi, 2002
Comments: Inhabits fast to moderate flowing premontane and montane streams and small rivers with gravel bottom.  Most abundantly found in shallow rapids and riffles during summer and moves to deeper places in winter.  Larvae and juveniles live on the bottom in very shallow shoreline habitats and leave the shores for faster-flowing waters as they grow.  Feeds on a wide variety of benthic vertebrates and algae.  Spawns in fast-flowing water (0.8-1 m/s) of shallow riffles with depth 0.5-0.8 m (Ref. 59043).