Author: De Vis, 1884
Comments: Adults are found abundantly in faster flowing sections of rivers, including riffles, over gravel, cobble and rocky bottoms.  Sexual maturity is reached at about 13 m for males.  Spawning occurs from July to October.  Feed on a variety of macroinvertebrates, including crustaceans, aquatic and terrestrial insects, and algae (Ref. 44894).  Eggs are guarded and fanned by the male parent (Ref. 205).