Author: Pastana, Dagosta & Esguícero, 2017
Comments: Found where water is transparent and the river bottom is composed of stones, sand and a moderate amount of vegetal debris. The type locality which is a hydroelectric reservoir (PCH Divisa) built in 2010 in the Rio Formiga, occupy an area of 6·8 km2. Before the dam was built, this place would have resembled the adjacent up and downstream areas of the river, i.e. a relative narrow (10–15 m wide) and shallow (maximum depth approximately 1·8 m in some sites) river stretch, with rapid waters and rifles. Stomach contents revealed a diet based on filamentous algae (c. 10%), vascular plant tissue (c. 6%), microcrustaceans (c. 10%), Chironomidae larvae (c. 10%) and Ephemeroptera nymphs (c. 64%). It may be considered an omnivorous species based on the main food resources exploited, but may have tendency toward invertivory. Approximately 50% of the analysed specimens had parasitic isopods (Cymothoidae) associated with the tongue. Specimens with this parasites exhibited a bag-shaped lower jaw, tongue atrophy and softening of the cartilaginous tissues (Ref. 128641).