Author: Boeseman, 1968
Comments: A rheophilic species, inhabiting largest part of freshwaters, from headwaters to upstream of the lower falls, with rocky substrate consisting of boulders, stones, gravels and sand; the exposed 
  wet rocks are also often covered by the Podostemaceae <i>Mourera fluviatilis</i> and <i>Apinagia richardiana</i> (Ref. 90186).