Author: Kuznetsov, 1888
Comments: Inhabits coastal fresh and brackish waters with salinities up to 7 ppt, and river mouths.  Occurs in groups.  From about 6 mm SL feeds actively on planktonic crustaceans, at 16-40 mm on larger copepods and mysids, and from second year of life on <i>Knipowitschia longecaudata</i>, <i>Clupeonella cultriventis</i>, benthic worms, molluscs and insect larvae. Attains first sexual maturity at 1 year of age.  Spawns in at least saline part of Gulf of Taganrog in June-July.  Adults migrate to open sea soon after spawning.  Eggs are released over mud bottom.  Eggs hatch in 2 days and larvae first lie on bottom, moving to the surface at 4 days (Ref. 59043).