Author: Fang & Kullander, 2009
Comments: The type locality is a small forest river with clear, colorless and slow-flowing or stagnant water and with bottom made up of stones, gravel and rock (Ref. 81819).  During low water season, this forest river is reduced to a series of connected pools, up to 2 m wide, and not more than 1 m deep (Ref. 81819).