Author: (Bussing, 1989)
Remark: Reviewed by Bussing (1998: 364), with key, diagnosis, geographical distribution, and figures.
Comments: Inhabits waters of all current velocities, but abounds mostly in swampy waters choked with vegetation.  Prefers waters of reduced size with soft substrata, between 1 and 70 m elevation.  Feeds on a variety of terrestrial and aquatic insects and occasionally small fish (Ref. 36880).