Author: Britz, Ali & Raghavan, 2012
Comments: Found in shallow part of a medium-sized clear water stream, up to 200 m wide and up to at least 2 m deep with sand, gravel and stones as substrate. Its type series was collected from a shallow part of the river with a fast current over medium-sized rounded stones and sandy patches as substrate.  Caught only from sandy areas (Ref. 93057).