Author: Karaman, 1928
Comments: Found along near shore of Lake Doiran.  Spawns in surf zone of the lake.  Pumping of water in the lake for irrigation has lowered its water level by at least 4 m since 1990 and has caused in the disappearance of subaquatic vegetation, leading to a severe ecological stress throughout the lake.  If pumping continues at its present rate, there is a risk that the lake will run dry (Ref. 59043). (Ref. 59043).