Author: Tan, 2020
Comments: This species occurs in lowland freshwater acid swamp-forest stream habitats, usually flowing into riverine habitats. Associated species include the following:  <i>Notopterus notopterus</i> (Notopteridae); <i>Barbodes cf. binotatus, Desmopuntius hexazona, Osteochilus vittatus, O. waandersii</i> (Cyprinidae); <i>Boraras maculatus, B. urophthalmoides, Rasbora bankanensis, R. cephalotaenia, R. usonensis, R. einthovenii, R. paucisqualis, Trigonopoma gracile, T. pauciperforatum</i> (Danionidae); <i>Acanthopsoides</i> sp., <i>Lepidocephalichthys furcatus, Pangio cuneovirgata, P. piperata, P. semicincta, P. muraeniformis</i> (Cobitidae); <i>Homalopteroides nebulosus</i> (Balitoridae); <i>Nemacheilus selangoricus</i> (Nemacheilidae); <i>Barbucca diabolica</i> (Barbuccidae); <i>Hemibagrus capitulum, Nanobagrus fuscus, Pseudomystus stenomus</i> (Bagridae); <i>Kryptopterus minor, Silurichthys hasseltii</i> (Siluridae); <i>Parakysis verrucosus</i> (Akysidae); <i>Clarias leiacanthus, C. meladerma</i> (Clariidae); <i>Monopterus javanensis</i> (Synbranchidae); <i>Neostethus smithi</i> (Phallostethidae); <i>Bihunichthys</i> sp., <i>Chaudhuria</i> sp. (Chaudhuriidae); <i>Oryzias minutulatus</i> (Adrianichthyidae); <i>Indostomus crocodilus</i> (Indostomidae); <i>Hemirhamphodon pogonognathus</i> (Zenarchopteridae); <i>Xenentodon canciloides</i> (Belonidae); <i>Doryichthys martensii</i> (Syngnathidae); <i>Brachygobius xanthomelas</i> (Gobiidae); <i>Pristolepis grooti</i> (Pristolepididae); <i>Nandus nebulosus</i> (Nandidae); <i>Betta imbellis, B. pi, stigmosa, Luciocephalus pulcher, Parosphomenus paludicola, Trichopodus trichopterus, Trichopsis vittata</i> (Osphronemidae); <i>Channa limbata</i>, and <i>C. lucius</i> (Channidae) (Ref. 122370).