Author: Near & Simmons, 2017
Comments: Habitats of this species differ between reproductive and non-reproductive periods. Both sexes in  in reproductive condition were most often found in swift runs of depths about one meter or less with predominant gravel and cobble substrates. On many occasions, adult males were collected in shallow, swift riffles with gravel bottoms during late April to early June. In summer months, adult males and females occurred in swift runs and in pool areas with substrates free of silt. Subadults were only encountered in pool habitats and were usually accompanied by adult females (Ref. 116752).