Author: Berg, 1910
Remark: This may represent two species (Ref. 59043).
Comments: Found in a wide range of cold and well oxygenated habitats from small, fast-flowing streams to large rivers.  Lives in groups.  Occurs in association with salmonid fishes or cyprinids of barbel zone.  Feeds on invertebrates, algae and detritus.  Spawns in shoals over clean gravel bottom in flowing water (Ref. 59043).