Author: Bohlen & Ŝlechtová, 2010
Comments: At the time of sampling, the type locality was a stream with width up to 8 m, but the riverbed would allow a 20 m wide river at high water level, and with calm stretches, depression up to 1 m deep as well as riffles. Riffle consisted of gravel with few larger stones and maximum water depth of 40 cm (Ref. 83421).