Author: Keith, Feunteun & Vigneux, 2010
Comments: Occurs in clear and calm streams with sandy or slightly rocky bottom and large pools.  Lives on the bottom of the river.  Also observed from sea level to 40 meters high in altitude.  Possibly spawns in freshwater, with free embryos drifting downstream to the sea where they undergo a planktonic stage, before returning to the rivers to grow and reproduce, hence are amphidromous (Ref. 83881).