Author: (Norman, 1926)
Remark: Redescription in Le Bail et al. (2000: 254).
Comments: Rarely attains a length of 20 cm (Ref. 27188).  Rheophilic species occurring in very sunny rapids with strong current and rocky bottom (Ref. 35381); inhabiting largest part of freshwaters, from headwaters to upstream of the lower falls, with rocky substrate consisting of boulders, stones, gravels, and sand; the exposed wet rocks are also often covered by the Podostemaceae <i>Mourera fluviatilis</i> and <i>Apinagia richardiana</i> (Ref. 90186).