Author: Kottelat, 1997
Comments: Lacustrine, in midwater. Spawns in December, along shores at 0-30 m depth. In both Lakes Thun and Brientz, there are forms (?) of this species spawning as deep as 70 m, some with a deeper body. They were reportedly more distinct earlier and had more distinct spawning seasons. The original pattern might have been blurred by hybridization with introduced species (Ref. 59043).