Author: Karaman, 1924
Comments: A short-lived species which occurs in lakes and water bodies on low-lying plains,  with little current.  Feeds on invertebrates and plants (Ref. 26100), preying mainly on <i>Gammarus</i> in lakes and on insects in streams (Ref. 59043).  Spawns in tributaries or on gravel beaches in lake, especially near underwater springs (Ref. 59043). Threatened by overfishing and water abstraction  (Ref. 26100).