Author: Watson & Pöllabauer, 1998
Comments: Prefers areas with gravel and overhanging vegetation in slower places of swift, clear streams; rocks and gravel usually coated with detritus.  Dives into the substrate when threatened, with its eyes, mouth and first dorsal fin above the gravel.  Feeds mainly on small freshwater crustaceans, especially transparent shrimp which are abundant. Ref. for amphidromous (Keith et al, in press 2014).