Author: (Wu & Ni, 1985)
Comments: Usually found in mangroves, at sites not far from the sea; collected by beam trawl in shallow water or by hand during low tide on mud-banks, in tiny puddles, on exposed mud surface or partly buried in mud.  One specimen in the Gulf of Carpentaria was found alive among the bases of dead mangrove <i>Bruguiera exaristata</i> on quite dry, firm mud (Ref. 80545).