Author: Stiassny, Rodriguez & Loiselle, 2002
Comments: Collected in the upper and middle reaches of the Lokoho river in the general vicinity of the town of Andapa.  Found predominantly in clear streams over gravel or coarse sand bottoms.  This streams are characterized by soft, slightly acidic to moderately alkaline water; total and carbonate hardness values < 1°DH and pH 6.8-7.5.  Collected from both rapidly flowing rivers and small brooks.  Stomach contents almost exclusively of macerated insect remains, most of terrestrial origin.  A few remains of aquatic insect larvae suggesting a somewhat opportunistic feeding (Ref. 42716).