Author: Naseka & Freyhof, 2004
Comments: Found in turbid waters with very strong current (Ref. 52363).  Occurs in large, high-gradient river stretches with fast but not turbulent current and gravel or sand bottom.  Active at night where it stays near shore on mud or sand bottom.  Feeds on benthic invertebrates, mainly Trichoptera larvae.  Appears to spawn late and to have a long spawning period.  Many small juveniles are observed in July (Ref. 59043).