Author: Berg, 1914
FBname: Kuban's nase
Comments: Inhabits piedmont and montane rivers with strong current and gravel bottom.  Occurs most of the year in the middle or upper stretches of rivers and moves downstream in winter to lower stretches or larger tributaries.  Feeds predominantly on detritus, but also takes periphyton, benthic invertebrates, especially during first year of life.  Spawns at sites with gravel or hard sand,  with rapids and swift current (up to 1.5 m/s), 0.2-1.0 m deep, in clean water with high oxygen concentrations.  Very sensitive to water quality, especially turbidity (Ref. 59043).