Author: (Greenwood & Kullander, 1994)
Comments: Rheophilic (Ref. 52307). Collected from clear, fast-flowing waters. The type series was taken from down to 2 m depth close to the shore, over rocky bottoms. Water temperature was 20.7 °C, pH 6.8, conductivity 25.7 µS, dissolved oxygen 9.3 mg/l (Ref. 26626).