Author: Thomas, Rodriguez, Carvallaro, Froehlich & Macedo Corrêa e Castro, 2013
Comments: Occurs in flowing water habitats ranging from small, seasonally intermittent streams (arroyos) to large permanent rivers.  Collected from the following localities:  repeatedly in small order streams with clear water in the upper rio Miranda drainage within the boundaries of the Parque Nacional da Serra da Bodoquena;  in and above riffles at depths of 0.3-1.5 m in the Aquidaban and Tebicuary drainages; in black water at the edge of the main river channel in swift current over sand and mud substrates in the rio Negro at the southern end of the Pantanal wetlands; along the shore over substrates of sand, silt and mud, at a depth of less than 1.5 m using seine at night in the lower rio Paraná (Ref. 94550).