Author: Li & Arai, 2014
Comments: The type locality is a hill stream with bottom consisting of mud mixed with stones, low water level, and depth mostly less than 100 cm. Collected from the upper portion of the stream.  Spawns in summer from June to August, several times during the spawning periods, and usually releases 10-20 eggs at one time.  Lives in host mussel <i>Ptychorhynchus murinum (Ref. 95613).