Author: Fowler, 1943
Remark: The type locality given by Fowler (1943) is not in accordance with the original label of the holotype (M.H. Sabaj, pers.comm.) or the information given by Brown (1941: 834) on the position of the Hacienda Mascota (see also Böhlke, 1958: 24-25). Type locality should be ammended to "Ecuador, Napo-Paztaza, Hacienda Mascota, río Topo, a tributary of río Paztaza, 1°25’S, 78°11’W."