Author: Allen & Unmack, 2008
Comments: Inhabits narrow (to about 15-20 m wide), clear rivers with gradual gradients flowing initially through second growth forest, but entering primary, nearly closed canopy forest after about 1 km upstream from the sea. Collected from deeper pools (to about 2-3 m) over limestone cobble/-gravel bottoms with slow to moderate flow rates; temperature of 25.5° and pH 8.7 (Ref. 74958).