Author: Trewavas, 1935
Comments: Is usually solitary over rocky substrates (Ref. 6307).  Feeds by nipping at the mat of epilithic algae.  The diet consists mainly of filamentous algae of the genus <i>Calothrix</i> and not of small invertebrates as is generally assumed.  Chironomid and centropogomid larvae, mites, ostracods and other small invertebrates comprise a small proportion of the gut contents (Ref. 4889).