Author: Suzuki, Oseko, Kimura & Shibukawa, 2020
Comments: Occurs in the upper reaches of swift freshwater streams in montane areas of Ryukyu Is., Japan. It appears to be amphidromous, judging from small-sized eggs laid by the females at the spawning grounds in Urauchi-gawa River, Iriomote-jima I. (Ref. 123096).