Author: Alonso, Terán, Aguilera, Říčan, Casciotta, Serra, Almirón, Benítez et al., 2019
Remark: Entered from e-publication.
Comments: Inhabits streams with sandy and rocky bottom and abundant marginal vegetation. The climate where it is found, on the mountain ridge in Aristóbulo del Valle, is classified as warm and temperate. In this area there is a significant rainfall throughout the year (average per year is 1,678 mm), and even in August with the lowest precipitation, it still has a considerable amount of rain (111 mm). The lowest precipitation occur from July to September, the driest period and lowest water levels from December to February due to the marked seasonal change in temperature, and peaks in precipitation occur from May and October. The annual average temperature is 19.1°C, with 23.9ºC monthly average temperature in January and 14.3ºC in June. Water level in the stream can have important changes: water turbidity is variable among the year with peaks of turbidity after rains and increases in water visibility the rest of the year. Specimens are frequently collected in the rocky areas or associated with marsh and marginal vegetation. The holotype was collected together with the female paratype under the marginal vegetation and with fry on November 2016 (Ref. 119666).