Author: Stiassny, Chakrabarty & Loiselle, 2001
Comments: Found in flowing water with isolated pools; rocky bottom interspersed with patches of cobble and coarse gravel (Ref. 40691). A biparental substratum-spawner that practices long-term care of its fry; breeding pairs guarding fry (Ref. 40691). Stomach contents were mainly crushed insects, most of which appear to be nymphal odonates and hemipterans, and significant amounts of fibrous plant material (Ref. 40691).