Author: (Nagelkerke & Sibbing, 1997)
Comments: Has a small preference for rocky substrates and waters less than 6 m deep.  Is a partial piscivore:  specimens smaller than 15 FL mainly eat insect larvae and zooplankton, but also some fish.  With size, the amount of fish in the diet increases, as does the amount of plant material, while the amount of insects decreases and zooplankton disappears altogether (Ref. 35835).