Author: Johnson & Randall, 2008
Comments: Collected on a broad shallow bank, on coarse sand and gravel substrate, which included numerous fallen leaves and the open shells of freshwater bivalve molluscs. The riverine environment at the type locality is flowing, clear and fresh (being about 5-6 km upstream from the brackish zone), but is clearly subject to tidal influence, with about a 75 cm rise and fall.  Environmental parameters at the time of collection (25 November 2006):  conductivity 59.6-60.6 μS/cm (approximately 30 ppm), pH 6.8, temperature 29°C, dissolved oxygen 6.5 ppm, and turbidity 0.8 NTU.  Stomach contents included fragments of insect larvae and numerous sand grains (Ref. 86741).