Author: (Günther, 1903)
Comments: Mainly herbivorous, but also takes small amounts of crustaceans and insects (Ref. 52307). Oviparous (Ref. 205). Pair-bonding, open (Ref. 52307) substrate brooder/spawner, with both parents guarding the brood (Ref. 52307, 55659). Eggs are laid and incubated in nests (Ref. 55659).