Author: Berg, 1948
Comments: Inhabits mountain lake (Lake Kezenoi-am) with altitude of 1870 m above sea level, with a maximum depth of 74 m, temperature about 5° C below 20 m and 5-18° C in upper layers in summer, covered by ice in winter, high oxygen concentrations down to bottom throughout the year and low plankton density.  This was the only fish inhabiting the lake prior to the introduction of <i>Gobio holurus</i> and <i>Squalius cephalus</i>.  Young juveniles feed predominantly on gammarids, chironomid larvae; larger juveniles and adults prey on molluscs, benthic invertebrates and fry; largest individuals are mostly piscivorous, especially since the introduction of <i>Gobio holurus</i>.  Spawns in lake, close to underwater springs.  Large-sized individuals probably undertake migration to tributaries.  Spawning extends over almost entire year.  Threatened due to the introduction of <i>Squalius cephalus</i>, which feeds on its fry (Ref. 59043).