Author: Fernholm, Norén, Kullander, Quattrini, Zintzen, Roberts, Mok & Kuo, 2013
Comments: The holotype was collected from the type locality over green muddy bottom at bottom temperature of 6.8°C, surface water temperature 28.3°C (Anonymous, 1910).   Additional specimens from the Verde Island Passage were collected at 578 and 710 m, sediment type ranged from mud to sand (Ref. 95645).