Author: (Dunz & Schliewen, 2010)
Comments: Deepwater specimens primarily feed on planktonic organisms in the open water column; inshore specimens pick on small particles from substrate and feed on allochtonous matter from water surface (Ref. 85854). The 'little-black' form breeds in all depth zones; the 'large-black' form breeds preferentially in log-holes of dead wood in the shallow region above 1 m (Ref. 85854).