Author: Shidlovskiy, Watters & Wildekamp, 2010
Comments: Found in ephemeral pools and swamps on floodplains; water level in the habitats is subject to seasonal changes and generally the water disappears completely during the dry season (Ref. 85866). Grass vegetation is usually found near the habitat margins; much of the swamps may be utilised by local inhabitants for the cultivation of rice; aquatic vegetation may comprise <i>Nymphea</i>, <i>Ottelia</i>, <i>Lagarosiphon</i> and <i>Utricularia</i> species (Ref. 85866). It has an annual mode of reproduction (Ref. 85866).