Author: Kullander, 2012
Comments: The type locality was a very small stream, about 50 cm wide, almost dry and with no other fish present.  Also found in a small stream, about 2 m wide and less than 1 m deep, with fast flowing cold water, no aquatic vegetation, lots of boulders and wood in the stream, substrate composed of sand and gravel. Another collection site was a stream up to 4 m wide and 2 m deep, with clear cool water, faster flowing parts but also deeper pools with slower water and no aquatic vegetation (Ref. 93124).