Author: Hosoya, Ito & Miyazaki, 2018
Comments: Occurs at the uppermost reaches of mountain streams surrounded by dense natural vegetation; preferring sandy bottoms with gravel or pebbles, where it hides under rocks and stones. It is syntopically distributed together with <i>L. echigonia</i> in the Yura and Kako rivers, although <i>L. echigonia</i> has a tendency to inhabit more lentic waters in swampy streams and ditches (Ref. 128930).