Author: Muller, Rapp Py-Daniel & Zuanon, 1994
Remark: 12S and 16S mitochondrial rRNA gene sequences used in molecular phylogenies by Montoya-Burgos et al. (1997: fig. 4; 1998: figs. 3 & 6; EMBL/GenBank Y08280; Y08328).
Comments: Inhabits clear running water, without a strong current to a depth of 3 meters during dry season.  Prefers narrow cracks of submerged rocks, small passages between superimposed rocks, or lives below flat rocks on the bottom.