Author: Takahashi & Hori, 2006
Comments: Occurs exclusively over rocky substratum, 1-5 m above the bottom from 5 m to at least 35 m depth; most abundant at ca. 15 m depth; females and sub-adult males forms mixed-species schools; adult males each maintain territories above adjacent large rocks or boulders; feed almost exclusively on cyploploid copepodids; mouthbrooding females (Ref. 57033). 