Author: (Nagelkerke & Sibbing, 1997)
Comments: Occurs over muddy, sandy and rocky substrates, preferably in water deeper than 6 m.  A real benthivore:  specimens smaller than 10 cm FL mainly eat insect larvae, but also zooplankton and ostracods.  With increased size, adult insects, mollusks and detritus also become important (Ref. 35835).