Author: (Lippitsch & Bouton, 1998)
Comments: Inhabits various stony and rocky habitats, including stone-sand mixed substrates.  It is generally more abundant at gently sloping shores with small and medium sized rock boulders.  Females, non-territorial males and subadults are gregarious and forage in groups.  Feeds on a variety of food items such as diatoms, filamentous epilithic algae, detritus, insect larvae and Bryozoa.