Author: Pereira, Vieira & Reis, 2007
Comments: The type locality is a small, shallow river with very clear, transparent water, slow to moderate current, and with bottom consisting of rocks, loose stones and sometimes gravel (Ref. 75793).  Collected in areas of rapids among loose stones and pebbles.  Nuptial males and larger individuals occur among larger stones and on the stronger current (Ref. 75793).