Author: (Fang & Kottelat, 1999)
Comments: Found in pools in small forest streams, sometimes even on remnant pools on very steep slopes (about 35°) and also in a mainstream at localities with various gradients, from standing to relatively swift waters.  All these habitats have clear to very clear water and forest cover (except downstream of the falls where during the dry season, most of the stream is covered by large basalt blocks or disappears in underground channels in the basalt).