Author: (Peters, 1868)
Comments: Collected from the lower freshwater reaches of a stream in Japan,150 m from the upper limit of tidal fluctuations, with <i>S. ninja</i>. It was also observed in two other streams on Okinawa Island, located 50 and 600 m from the upper limit of tidal fluctuations. Found in rapids at depths of 5-30 cm and in one case, was observed to swim away in the middle of the water column against a strong current,  a behavior not seen in <i>S. ninja</i> and <i>S. ampluvinculus</i> (Ref. 125666).